aTYR PHARMA INC (ATYR) — Institutional Holders, Q2 2026

90 institutional investment managers reported a ATYR position on SEC Form 13F-HR for the quarter ended June 30, 2026, with a combined reported value of $23.2M. Every filer is listed below, ranked by position value. Put and call option positions are excluded, and a firm filing several lots or share classes is shown once with its total.

← ATYR stock profile

ATYR in Q2 2026: what funds did

Funds that filed a 13F for both Q1 2026 and Q2 2026, by what they did with the common stock. Share counts are adjusted for any split between the two quarters; dollar figures are the change in shares at this quarter’s price.

New positions

Added to

Trimmed

Sold out

Biggest buyers

AQR CAPITAL MANAGEMENTAdded · +2,114,530 sh+$1.3M
Kalehua Capital ManagementNew · +1,788,167 sh+$1.1M
GSA CAPITAL PARTNERS LLPAdded · +1,770,277 sh+$1.1M
JANE STREET GROUPAdded · +1,672,795 sh+$998.3K
CITADEL ADVISORSAdded · +1,194,695 sh+$713.0K

Biggest sellers

FMRTrimmed · -12,235,425 sh-$7.3M
BlackRockTrimmed · -4,616,572 sh-$2.8M
UBS Group AGTrimmed · -2,336,062 sh-$1.4M
FEDERATED HERMESTrimmed · -2,000,000 sh-$1.2M
STATE STREETTrimmed · -1,259,448 sh-$751.6K

ATYR 13F holders ranked 1–90 of 90

Institutional investors reporting a position in aTYR PHARMA INC on SEC Form 13F-HR for Q2 2026, ranked by reported value.
#	Institution	Shares	Value	% of portfolio	QoQShares vs Q1 2026
1	FEDERATED HERMES#1 · 8,509,678 sh	8,509,678	$5.1M	<0.01%	-19.0%
2	VANGUARD CAPITAL MANAGEMENT#2 · 3,808,091 sh	3,808,091	$2.3M	0.00%	-0.1%
3	GSA CAPITAL PARTNERS LLP#3 · 3,454,863 sh	3,454,863	$2.1M	0.13%	+105.1%
4	AQR CAPITAL MANAGEMENT#4 · 2,610,548 sh	2,610,548	$1.6M	<0.01%	+426.3%
5	FMR#5 · 2,471,418 sh	2,471,418	$1.5M	<0.01%	-83.2%
6	BlackRock#6 · 1,827,332 sh	1,827,332	$1.1M	0.00%	-71.6%
7	Kalehua Capital Management#7 · 1,788,167 sh	1,788,167	$1.1M	0.76%	New
8	JANE STREET GROUP#8 · 1,721,297 sh	1,721,297	$1.0M	<0.01%	+3448.9%
9	UBS Group AG#9 · 1,531,151 sh	1,531,151	$914.1K	<0.01%	-60.4%
10	MORGAN STANLEY#10 · 1,343,636 sh	1,343,636	$801.9K	0.00%	+9.6%
11	TWO SIGMA INVESTMENTS#11 · 1,245,147 sh	1,245,147	$743.1K	<0.01%	+8.8%
12	CITADEL ADVISORS#12 · 1,242,067 sh	1,242,067	$741.3K	<0.01%	+2521.9%
13	GEODE CAPITAL MANAGEMENT#13 · 1,177,020 sh	1,177,020	$702.7K	0.00%	-50.7%
14	SUSQUEHANNA INTERNATIONAL GROUP, LLP#14 · 747,039 sh	747,039	$445.8K	0.00%	-13.8%
15	MILLENNIUM MANAGEMENT#15 · 656,051 sh	656,051	$391.5K	<0.01%	-60.0%
16	GOLDMAN SACHS GROUP#16 · 641,208 sh	641,208	$382.7K	0.00%	+126.4%
17	VANGUARD FIDUCIARY#17 · 576,125 sh	576,125	$343.9K	<0.01%	-11.6%
18	STATE STREET#18 · 494,951 sh	494,951	$295.4K	0.00%	-71.8%
19	MARSHALL WACE, LLP#19 · 368,120 sh	368,120	$220.9K	<0.01%	+166.5%
20	XTX Topco Ltd#20 · 226,678 sh	226,678	$135.3K	<0.01%	+103.4%
21	NORTHERN#21 · 220,141 sh	220,141	$131.4K	0.00%	-69.0%
22	BANK OF AMERICA#22 · 207,468 sh	207,468	$123.7K	0.00%	-31.7%
23	RENAISSANCE TECHNOLOGIES#23 · 165,222 sh	165,222	$98.6K	<0.01%	+307.7%
24	JPMORGAN CHASE#24 · 150,666 sh	150,666	$85.3K	0.00%	+258.8%
25	GROUP ONE TRADING, L.P.#25 · 138,878 sh	138,878	$82.9K	<0.01%	-24.5%
26	Qube Research & Technologies Ltd#26 · 126,996 sh	126,996	$75.8K	<0.01%	New
27	HRT FINANCIAL#27 · 126,681 sh	126,681	$75.0K	<0.01%	New
28	DIMENSIONAL FUND ADVISORS#28 · 90,861 sh	90,861	$54.1K	0.00%	-1.1%
29	HSBC PLC#29 · 66,185 sh	66,185	$38.4K	0.00%	+263.4%
30	Mariner#30 · 58,658 sh	58,658	$35.0K	0.00%	New
31	HOHIMER WEALTH MANAGEMENT#31 · 55,000 sh	55,000	$32.8K	<0.01%	-45.0%
32	MERCER GLOBAL ADVISORS#32 · 53,531 sh	53,531	$31.9K	0.00%	+252.1%
33	KINGSWOOD WEALTH ADVISORS#33 · 52,000 sh	52,000	$31.0K	—	0.0%
34	Schonfeld Strategic Advisors#34 · 50,880 sh	50,880	$30.4K	<0.01%	New
35	ADAR1 Capital Management#35 · 50,685 sh	50,685	$30.2K	<0.01%	0.0%
36	Alpha Wealth Funds#36 · 50,000 sh	50,000	$29.9K	0.02%	New
37	Tower Research Capital (TRC)#37 · 48,768 sh	48,768	$29.1K	<0.01%	+179.9%
38	Virtu Financial#38 · 46,921 sh	46,921	$28.0K	<0.01%	+17.2%
39	Creative Planning#39 · 41,593 sh	41,593	$24.8K	0.00%	+295.8%
40	WEALTH ENHANCEMENT ADVISORY SERVICES#40 · 43,948 sh	43,948	$23.9K	—	+182.1%
41	Point72 Asset Management, L.P.#41 · 37,278 sh	37,278	$22.2K	0.00%	New
42	Hobbs Group Advisors#42 · 35,992 sh	35,992	$21.5K	<0.01%	+85.3%
43	LPL Financial#43 · 35,549 sh	35,549	$21.2K	0.00%	0.0%
44	CAPTRUST FINANCIAL ADVISORS#44 · 33,946 sh	33,946	$20.3K	0.00%	+112.4%
45	Vanguard Global Advisers#45 · 33,414 sh	33,414	$19.9K	0.00%	0.0%
46	BARCLAYS PLC#46 · 31,029 sh	31,029	$18.5K	0.00%	-31.2%
47	RHUMBLINE ADVISERS#47 · 30,412 sh	30,412	$18.2K	0.00%	-70.5%
48	Cerity Partners#48 · 29,232 sh	29,232	$17.4K	0.00%	New
49	Rockefeller Capital Management L.P.#49 · 28,118 sh	28,118	$16.8K	0.00%	0.0%
50	BNP PARIBAS FINANCIAL MARKETS#50 · 25,055 sh	25,055	$15.0K	0.00%	-0.9%
51	BALYASNY ASSET MANAGEMENT L.P.#51 · 23,015 sh	23,015	$13.7K	0.00%	+122.8%
52	Y-Intercept (Hong Kong) Ltd#52 · 22,373 sh	22,373	$13.4K	—	New
53	Abel Hall#53 · 22,117 sh	22,117	$13.2K	—	New
54	Steward Partners Investment Advisory#54 · 19,600 sh	19,600	$11.7K	<0.01%	+18.1%
55	MAI Capital Management#55 · 16,846 sh	16,846	$10.1K	0.00%	+24.7%
56	RAYMOND JAMES FINANCIAL#56 · 16,801 sh	16,801	$10.0K	0.00%	+1.2%
57	OF TOLEDO NA#57 · 16,640 sh	16,640	$9.9K	—	0.0%
58	SRS Capital Advisors#58 · 16,557 sh	16,557	$9.9K	<0.01%	New
59	Vanguard Personalized Indexing Management#59 · 14,899 sh	14,899	$8.9K	<0.01%	-10.6%
60	Close Asset Management Ltd#60 · 12,300 sh	12,300	$7.0K	<0.01%	-24.1%
61	BRIGHTON JONES#61 · 11,458 sh	11,458	$6.8K	<0.01%	New
62	Gladstone Institutional Advisory#62 · 11,300 sh	11,300	$6.7K	<0.01%	0.0%
63	WELLS FARGO & COMPANY#63 · 6,500 sh	6,500	$3.9K	0.00%	-86.0%
64	Integrated Wealth Concepts#64 · 5,995 sh	5,995	$3.6K	0.00%	New
65	ROYAL BANK OF CANADA#65 · 5,996 sh	5,996	$3.0K	0.00%	-46.2%
66	Allworth Financial#66 · 3,615 sh	3,615	$2.2K	0.00%	+887.7%
67	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AME#67 · 2,011 sh	2,011	$1.2K	0.00%	0.0%
68	Ares Financial Consulting#68 · 2,000 sh	2,000	$1.2K	—	0.0%
69	Newbridge Financial Services Group#69 · 1,979 sh	1,979	$1.2K	—	-4.8%
70	Vision Retirement#70 · 1,500 sh	1,500	$895.00	<0.01%	0.0%
71	MCF Advisors#71 · 1,400 sh	1,400	$836.00	0.00%	+250.0%
72	JONES FINANCIAL COMPANIES LLLP#72 · 1,500 sh	1,500	$791.00	0.00%	0.0%
73	PNC FINANCIAL SERVICES GROUP#73 · 1,276 sh	1,276	$762.00	0.00%	-84.4%
74	EverSource Wealth Advisors#74 · 1,149 sh	1,149	$685.00	0.00%	-99.8%
75	CITIGROUP#75 · 900 sh	900	$537.00	0.00%	-84.7%
76	ROTHSCHILD INVESTMENT#76 · 885 sh	885	$528.00	0.00%	+208.4%
77	Clearstead Advisors#77 · 861 sh	861	$515.00	0.00%	New
78	Essential Partners#78 · 815 sh	815	$486.00	—	-49.6%
79	GAMMA Investing#79 · 659 sh	659	$393.00	—	-91.5%
80	True Wealth Design#80 · 464 sh	464	$277.00	—	+544.4%
81	Farther Finance Advisors#81 · 372 sh	372	$222.00	—	-71.8%
82	SJS Investment Consulting#82 · 208 sh	208	$125.00	—	+395.2%
83	Larson Financial Group#83 · 150 sh	150	$90.00	0.00%	0.0%
84	IFP Advisors#84 · 143 sh	143	$85.00	—	-97.6%
85	CALDWELL SUTTER CAPITAL#85 · 52 sh	52	$31.00	0.00%	0.0%
86	IFG Advisory#86 · 50 sh	50	$30.00	0.00%	New
87	NORTHWESTERN MUTUAL WEALTH MANAGEMENT#87 · 25 sh	25	$15.00	0.00%	New
88	NISA INVESTMENT ADVISORS#88 · 8 sh	8	$5.00	—	-99.2%
89	Caitong International Asset Management , Ltd#89 · 1 sh	1	$1.00	0.00%	0.0%
90	FNY Investment Advisers#90 · 1,500 sh	1,500	$0.00	—	0.0%

Funds that sold out of ATYR in Q2 2026 (35)

Held ATYR at the end of Q1 2026 and filed a Q2 2026 13F without it. Not in the list above, because they no longer hold it. Largest prior position first.

Institutions that held ATYR in Q1 2026 and reported no position in Q2 2026.
Institution	Shares sold	Value in Q1 2026
VANGUARD PORTFOLIO MANAGEMENT569,227 sh	569,227	$444.0K
Nuveen229,284 sh	229,284	$178.8K
CHARLES SCHWAB INVESTMENT MANAGEMENT210,737 sh	210,737	$164.4K
Bank of New York Mellon193,531 sh	193,531	$151.0K
DEUTSCHE BANK AG\103,326 sh	103,326	$80.6K
VICTORY CAPITAL MANAGEMENT88,436 sh	88,436	$69.0K
ALLIANCEBERNSTEIN L.P.64,480 sh	64,480	$50.5K
Redhawk Wealth Advisors49,150 sh	49,150	$38.3K
Invesco Ltd.46,354 sh	46,354	$36.2K
PRICE T ROWE ASSOCIATES46,126 sh	46,126	$36.0K
MIRAE ASSET GLOBAL ETFS Ltd.46,073 sh	46,073	$35.9K
Belpointe Asset Management45,738 sh	45,738	$35.7K
SEI INVESTMENTS33,829 sh	33,829	$26.4K
NEW YORK STATE COMMON RETIREMENT FUND27,500 sh	27,500	$21.4K
INTECH INVESTMENT MANAGEMENT26,187 sh	26,187	$20.4K
Police & Firemen's Retirement System of New Jersey26,081 sh	26,081	$20.3K
Neuberger Berman Group21,743 sh	21,743	$17.0K
RBF Capital20,000 sh	20,000	$15.6K
PRUDENTIAL FINANCIAL14,660 sh	14,660	$11.4K
ProShare Advisors13,905 sh	13,905	$10.8K
Crossmark Global12,666 sh	12,666	$9.9K
Pasadena Private Wealth12,228 sh	12,228	$9.5K
Apollon Wealth Management11,900 sh	11,900	$9.3K
Russell Investments Group, Ltd.11,361 sh	11,361	$8.9K
MANUFACTURERS LIFE INSURANCE COMPANY, THE10,552 sh	10,552	$8.2K
VOYA INVESTMENT MANAGEMENT10,338 sh	10,338	$8.1K
Ameritas Investment Partners8,881 sh	8,881	$6.9K
Legal & General Group Plc7,920 sh	7,920	$6.2K
OSAIC5,000 sh	5,000	$3.9K
TOTH FINANCIAL ADVISORY1,400 sh	1,400	$1.1K
SIMPLEX TRADING1,335 sh	1,335	$1.0K
Sterling Capital Management742 sh	742	$579.00
Wealth Preservation Advisors157 sh	157	$122.00
Root Financial Partners9 sh	9	$7.00
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM5,190 sh	5,190	—

Source: Form 13F-HR filings on SEC EDGAR, Q2 2026. Values are as reported by each manager at quarter end. “% of portfolio” is the position’s share of the manager’s total reported 13F value for the same quarter. “New” means the manager filed a 13F for Q1 2026 without this position; a dash means no comparison is available. Where a manager’s own filing is internally inconsistent its value is withheld and it is listed last. 13F data shows reported holdings, not recommendations.